This is a common heightmap editor bug. When you save the heightmap, it will create a hole in the corner of each quad where elevation drops to 0 or a spike where it goes up to the max elevation (every 32 cells, so at 0,0, at 31,0, etc.).
To fix it, load your worldspace in the cell view window at these cells and use the landscaping tool to flatten out the hole. If you save your mod without correcting these errors and load it up in-game the landscape mesh in these cells will be missing (but the objects you place will still be there). In this case, just correct your land mesh in the cell view window using the landscape editor as described and your mod will be fine. Fixing it in the heightmap editor won't solve the problem because that is the tool causing the problem.
It seems like a hassle at first, but it only takes a few seconds to fix the holes and there aren't many of them. You just have to do it each time you save the heightmap.